The Cerritos crew's mission to deliver the U.S.S. Voyager to Earth to be a museum is interrupted when a transporter accident merges Dr. T'Ana and chief engineer Billups into a single life form.
Mariner resolves to get off Ransom's good side by screwing up a mission to an alien menagerie. Rutherford tries to earn a quick promotion, and Boimler's new quarters prove to be not all they're cracked up to be.
The crew visits a ring world in crisis after its artificial intelligence goes on the fritz. Boimler leads his first away mission, and the other lower deckers suspect they are being hazed following their recent promotions.
Mariner and T'lyn accompany Tendi to the Orion homeworld to visit her family, where they learn that her sister has been kidnapped on the eve of her wedding. Boimler and Rutherford look for a way to resolve their differences.
The Cerritos pays a visit to the Ferengi homeworld after the Grand Nagus makes an overture to join the Federation. While Tendi and Rutherford pretend to be a married couple, Mariner is reunited with an old friend.
After Badgey attacks the Cerritos Rutherford tries to stop his creation's megalomaniacal takeover of the quadrant. Meanwhile Boimler and Tendi find themselves in the middle of AGIMUS and Peanut Hamper's attempt to escape from sentient A.I. prison.
Three computerized villains return to cause problems for the Cerritos crew. (Paramount+)
The Lower Deckers are trapped when another standard cave mission goes awry, requiring them to rely on the skills they learned other times they were trapped in caves.
Boimler, Tendi, and T'lyn try to keep Mariner out of danger, only for the away team to find themselves stranded on a planet with castaways from various species. Meanwhile Captain Freeman leads a team to a black market bar to obtain intel on a Federation citizen who may have gone rogue.
Mariner is reunited with someone from her past at Starfleet Academy, who has managed to assemble an interspecies fleet of down-and-out lower decks officers.